You may find this source more understandable and valid as set out by Stanley Coren. Lee Charles Kelley's writings are also a good resource. I can spend days in Psychology Today's Animal Behavior section.
GSD ~ Fortune zu Treuen Handen "Frank" born 6/19/13
“The saddest aspect of life right now is that science gathers knowledge faster than society gathers wisdom.” ― Isaac Asimov